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The morphology and taxonomy is discussed of the Polypodiaceous genera Selliguea, Crypsinus, Holcosorus, Phymatopsis, Grammatopteris, Pycnoloma, Oleandropsis and Crypsinopsis, based on a revision of species from Malesia and the Pacific. All these genera are merged in Selliguea. Fifty-two species are revised, 32 new combinations are made. See also the description of a new species on p. 108, added after the proof stage.
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The genus Lepisorus J.Sm. is represented in Malesia with four species. One new species and one new combination are described here.
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The new combination Cadetia maliliensis (J.J. Sm.) Schuit. is made, based on Dendrobium mali-liense J.J. Sm. The genus Cadetia is recorded from Borneo for the first time.
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A new species is described in Tetrapetalum, a small genus of which two species have been known. Although Van Heusden (1992) stated that Tetrapetalum differs from Cyathostemma only in its dimerous flowers, Tetrapetalum seems to be more closely related with Uvaria than with Cyathostemma, because of spreading petals and unlobed stigmas. A key to the three species of Tetrapetalum is provided here.
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Details of a palynological record of Ulmus in North Sumatra extending from c. 21,250 B.P. are presented and compared with recent findings from Java and Sulawesi (Table 1).
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Shirakia (Euphorbiaceae) was described by Hurusawa (1954) with three species. It was named after Croton shiraki, a nomen nudum used by Siebold & Zuccarini (1843) for the species currently known as S. japonica (Siebold & Zucc.) Hurus. Kruijt (1996) transferred five additional species to Shirakia. Esser et al. (1998) showed that S. japonica, the type, differs in several characters from the remainder of the genus. In a phylogenetic analysis they concluded that all additional species have to be removed to other genera (Shirakiopsis Esser ined., Triadica Lour.), with Shirakia remaining as a monotypic genus endemic to Japan and adjacent East Asia.
The name, however, is a later homonym of Shirakia S. Kawasaki, a fern fossil described in 1934 from South Korea. This information is included in the WWW version of the Index Nominum Genericorum (Farr & Zijlstra, 1996 and onward). Because this earlier homonym was published in a publication of very restricted distribution, it was overlooked by recent workers (Kruijt, 1996; Esser et al., 1998). Gea Zijlstra and John Wiersema kindly confirmed that a new name is still necessary for Shirakia Hurus., and this is provided herewith.
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A cladogram of the subtribe Lasiococcinae (Homonoia, 2 species, Lasiococca, 3 species, and Spathiostemon, 2 species) is presented with the genus Wetria as outgroup. All three taxa are monophyletic groups of species with Lasiococca and Spathiostemon as sistergroups and Homonoia related to both of them. Within Lasiococca, L. comberi and L. malaccensis are probably closest related.
The two species of Homonoia are rheophytes, one is restricted to India where it shows two distinct forms, the other species is widespread from India throughout Malesia.
Lasiococca is represented by one species in Malesia, L. malaccensis, only known from three localities, ranging from the Malay Peninsula to Sulawesi and the Lesser Sunda Islands.
Spathiostemon has two species in Malesia, one is widespread in Malesia, the other one is restricted to part of Peninsular Thailand.
Clonostylis, a monotypic genus only known from the Sumatran type specimen, is not synonymous with Spathiostemon. Clonostylis is seemingly most similar to Mallotus and Macaranga.
Ricinus, also a monotypic genus, is introduced to Malesia and is generally cultivated. It is not part of the Lasiococcinae. The presence of phalanged stamens, also typical for the Lasiococcinae, is a parallel development as Ricinus shows many more androphores and the connective is often appendaged. Ricinus cannot readily be classified and retaining it in its present monotypic subtribe seems to be the best solution.
Wetria shows two species in Malesia. One is restricted to West Malesia, the other one is found in New Guinea and appears to be similar to the recently described W. australiensis of Queensland.
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The phenomenon of myrmecosymbiosis in Myristica, only occurring in some species from New Guinea, is reviewed here. A key to myrmecophilous taxa (and resembling species) is presented, and the status of their myrmecophily is briefly discussed. Three myrmecophytes are described as new species (M. dasycarpa, M. sarcantha, M. verruculosa), and M. subcordata var. rimosa is described as a new variety in a species liable to be confused with a myrmecophyte.
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An account of native and introduced Epipremnum species in West and Central Malesia is presented as a precursor to the forthcoming Flora Malesiana Araceae treatment. Seven species and one cultivar, none new, are recognized. Epipremnum grandifolium is returned to Scindapsus, the genus in which it was originally described and Scindapsus dilaceratus (synonyms Monstera dilacerata, Tornelia dilacerata), a long-overlooked synonym of E. pinnatum, is discussed. Related genera and identification pit-falls are highlighted. Dichotomous keys to the genera of tribes Anadendreae and Monstereae and to West and Central Malesian Epipremnum species, a multi-access key to Epipremnum species in West and Central Malesia and a geographical species-finder list are provided. All names in the review area are accounted for. Five species are illustrated.
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A new species, Ceropegia andamanica, allied to C. metziana Miq., from the Mount Harriet National Park in South Andaman is described and illustrated. It is the first record of the genus Ceropegia from the Andaman and Nicobar Islands, and it is currently known from a few scattered patches on just one island in areas which have been cleared of their native vegetation.
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Sections and macerations of roots and stems of Orchidantha maxillarioides (Ridl.) K. Schum. (Lowiaceae) were examined with a scanning electron microscope (SEM). Vessels with long scalariform perforation plates occur in both roots and rhizome; perforation plates have more pit membrane remnants in rhizomes. At least a few tracheids are present. The degree of perforation plate specialization correlates with the moist forest understory habit of Orchidantha, and is similar to that in primitive families of monocotyledons that occupy marshy habitats. Vessel data suggest a primitive position for Lowiaceae within Zingiberales in accordance with recent phylogenies of monocotyledons, but recent molecular data point to a position more nearly terminal in the clade; this situation needs resolution. Raphides are figured with SEM, as are starch grains with rounded outlines, the shape of the latter newly reported for Lowiaceae.
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Of the family Euphorbiaceae three new species of Mallotus Lour. [section Hancea (Seem.) Pax & K. Hoffm.] from Malesia are described and illustrated. Mallotus cordatifolius and M. wenzelianus from the Philippines, and M. grandistipularis from Central Sumatra, Indonesia.
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The genera Dioclea Humb., Bonpl. & Kunth, Luzonia Elmer, and Macropsychanthus Harms are briefly discussed. Keys to the species and notes to various taxa are given. Dioclea decandra Amshoff is proposed as a new name for Macropsychanthus ferrugineus Merr. Macropsychanthus lauterbachii Harms is lectotypified; its subspecies glabricalyx Verdc. and parviflorus Verdc. are lowered in rank to varieties, subsp. neobrittanicus Verdc. is synonymized to var. parviflorus (Verdc.) Adema.
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New material of Knema and Myristica necessitated the description of some new species, subspecies and forms, viz. Knema korthalsii subsp. rimosa, K. kunstleri subsp. pseudostellata, K. latericia subsp. ridleyi forma nana and forma olivacea, K. minima, K. tridactyla subsp. pachydactyla and subsp. salicifolia, and Myristica inaequalis and M. rosselensis subsp. minutiflora. Additional comments to other taxa are given and, where appropriate, insertion in (earlier published) general and regional keys are proposed.
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Last modified August 24, 1998 by P. Hovenkamp.