Lithocarpus echinifer (Merr.) A.Camus, Bull. Soc. Bot. Fr. 80 (1934)
Latin for 'sea-urchin', referring to the spiny fruits.

Synonyms
Quercus echinifera Merr.

Description
Mid-canopy tree up to 35 m tall and 80 cm dbh. Stipules ca. 1 mm long. Leaves alternate, simple, penni-veined, glabrous. Flowers ca. 2 mm diameter, green-yellow, placed in racemes. Fruits ca. 38 mm diameter, greenish, nut alomost completely enclosed by spiny cupule, spines curved downwards.

Ecology
In undisturbed mixed dipterocarp and sub-montane forests up to 1800 m altitude. Usually along rivers on sandy to clay soils. In secondary forests usually present as a pre-disturbance remnant tree.

Distribution
Borneo (Sarawak, Brunei, Sabah, East-Kalimantan).