Malesian Euphorbiaceae Newsletter 3
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Changes in Malesian Euphorbiaceae Genus List
Genera and Species to be revised
Additions and Changes in the Mailing List
The third edition of the Malesian Euphorbiaceae Newsletter has been finished. It has been completely compiled again by myself. I like to emphasize that everybody is invited to send in short notes. I very much like to receive notes about progress, field trips, and other events. This way we can update each other about the state of affairs in the revision of the Malesian Euphorbiaceae.
The newsletter is becoming popular, the mailing list is still increasing. Also the number of participants is still increasing, only a few genera are left which are not yet reserved by a researcher for a revision.
In this edition of the newsletter I like to draw your attention to new literature, progress reports, and the Flora Malesiana Symposium
In the next issue special attention will be given to the Expert Center for Taxonomic Identification (ETI).
Recently, the first part of the proceedings of the International Conference on the Systematics of the Euphorbiaceae was published. The conference was held in St. Louis in August 1989. In this first part, of these very belated proceedings, Grady Webster presents a classification and synopsis of the Euphorbiaceae, containing keys to and descriptions of the infrafamily levels and the genera. Per taxon a full account of its synonymy is presented. The account does not as much provide a new critical delimitation within the Euphorbiaceae, but is more an overview of the presently (more or less) accepted genera. The family view is broad, most small split families have been included: Stilaginaceae (Antidesma), Pandaceae (Galearia, Microdesmis), and Bischofiaceae (Bischofia). Not included are the small families Buxaceae and Daphniphyllaceae. This concept of the Euphorbiaceae equals the one used for the genera list of Malesian Euphorbiaceae (see below). The genera list includes the Stilaginaceae, Pandaceae, and Bischofiaceae and excludes the Buxaceae and Daphniphyllaceae.
Webster's treatment incorporates a natural system, therefore, the keys follow the classification. For this reason, the keys may be important to identify Malesian specimens up to the genus level. The artificial key published by Airy Shaw (e.g., the issues on Bornean and New Guinean Euphorbiaceae) is difficult to use. Webster's key, with a completely different structure, might therefore be helpful. However, with both keys complete material (male and female flowers and fruits) is often still needed.
Other papers presented during the conference will be published in future issues of the Annals of the Missouri Botanical Garden.
Webster, G.L. 1994. Systematics of the Euphorbiaceae: Introduction. Ann. Missouri Bot. Gard. 81: 1,2.
Webster, G.L. 1994. Classification of the Euphorbiaceae. Ann. Missouri Bot. Gard. 81: 3-32.
Webster, G.L. 1994. Synopsis of the genera and suprageneric taxa of Euphorbiaceae. Ann. Missouri Bot. Gard. 81: 33--144.
Two small genera have been completed in the mean time, Chaetocarpus and Neoscortechinia. Especially the genus Neoscortechinia was difficult, the differences between the species are very small. Six species have been recognized, most of them are wide-spread in W Malesia. One variety will be raised to specific rank, this species occurs locally in NE Borneo. Another very wide-spread species will be split up into two, a W Malesian species and an E Malesian species (New Guinea and the Solomon Islands). The revision, which includes a phylogenetic analysis with Cheilosa montana as outgroup, will probably be published in the next issue of Blumea. Therefore, no names are presented here, otherwise the new combinations will have been made in this obscure 'journal'.
The genus Chaetocarpus, a pantropical genus, is only represented with one species in Malesia, Ch. castanocarpus (Roxb.) Thwaites (India to W Malesia). Two more species can be found in Sri Lanka (Ceylon). One of them, Ch. pubescens (Thwaites) Hook.f., resembles a specimen collected in Irian Jaya (W New Guinea). The latter specimen was sterile except for some fruit remnants in an attached paper bag. No taxonomic status has been given to this specimen. (Unfortunately, Irian Jaya is still a major undercollected area, but presently, two projects have started in the Bird's head of New Guinea, which might reveal more and better specimens.) The revision will probably be published in the Indian journal Rheedea. Ch. castanocarpus, although vegetatively not very attractive, has very special, beautiful fruits (stinging hairs) and flowers (stamens united in an androphore; stigma papillae dendritic).
Two weeks ago I received a letter from Prof. Nguyen Nghia Thin, that he has finished the revision of the genera Lasiococca and Strophioblachia. As soon as I have received his manuscripts extra attention will be given to his results. Hopefully, this might already be in the next issue of the newsletter.
In the former newsletter the grants of the European Community were already mentioned. Two postdoctoral fellowships within the 'Botanical Diversity of the Indo-Pacific region Human Capital and Mobility Programme' have been assigned to the revision of Euphorbiaceae. Mrs. Petra Hoffmann will start with the genus Antidesma in Kew, England, for the next two years and Stefan Dressler will work on the genera Bridelia and Cleistanthus in Leiden, The Netherlands. They are both from the Humboldt University in Berlin and they will start their new jobs on September, 1st. Petra Hoffmann will, after having finished Antidesma, finish the revision of Actephila.
Two more M.Sc. students in Bogor, Indonesia, have started a revision of Euphorbiaceae genera. Mrs. Musayyinah and Mrs. Endah Woro Wiryani will work on the genera Koilodepas and Codiaeum respectively. Their supervision, just like with the other three M.Sc. students, will be in the experienced hands of Dedy Darnaedi, Johanis Mogea, Mien Rifai, Rugayah, and Elisabeth Widjaja.
In Leiden we have also started the application procedure for a Ph.D. fellowship within the 'Pathways from Asia to New Guinea' programme of the Dutch Science Foundation (NWO). This Ph.D. fellowship will include the revision of Baccaurea. After the revision a phylogenetic analysis will follow and the resulting data will be combined with those of the two other Ph.D. students in the programme working on insects. Finally, a historical biogeographic analysis will have to show, which geographical patterns are involved in distributions from SE Asia to New Guinea.
The third Flora Malesiana Symposium will held from 10-14 July in Kew, England. Last week the second and final circular has been published, which includes the registration forms and the abstract forms. Copies can be obtained from the secretary of organising committee, Mr. B.J. Blewett (Royal Botanic Gardens, Kew, Richmond, Surrey, TW9 3AE, United Kingdom).
The circular also contains information for those seeking funds for their participation. The circular urges everybody to apply for grants in their own country, but three British organisations might also be helpful in providing special grants. The Kew herbarium can assist in writing invitation letters.
Programme:
Registration will be on Sunday, 9 July (followed by a dinner at St. Mary's University College), which will be continued on Monday morning.
On Monday the opening will be followed by a lecture of a/the keynote speaker and the current state of Flora Malesiana. In the afternoon there will be workshops and contributed papers.
Tuesday will start with phytogeography and cladistic studies, followed again in the afternoon by workshops and contributed papers.
On Wednesday local flora projects will present themselves and a poster session will be held. At the end of the afternoon there will be short excursions and the Flora Malesiana Board Meeting.
Monographs and biological papers will be presented on Thursday morning, while in the afternoon there will be a Computer applications workshop and contributed papers. The special Symposium Dinner, with an after dinner speech by Dr. Whitmore, will be in the evening.
Finally, on Friday, papers on ethnobotany, conservation, and ecology will be followed by summaries of the workshops and the closure of the symposium.
For those interested, the week after the Flora Malesiana Symposium, the Holttum Memorial Pteridophyte Symposium will be organised in Kew and for those staying over from the FM III a special excursion is organised on Sunday.
Papers and Posters
Everybody who likes to contribute a paper or a poster (display boards of 2 by 1 m are available) should send an outline for consideration to the Scientific Programme coordinator, Mr. M.J.E. Coode (address see above). A special committee 'will select and advise authors whether their presentations have been designated papers or posters'.
Herbarium visit
Those people who like to visit the Kew herbarium should, at least one month in advance, contact Mr. B.J. Blewett (address see above), and those who like to visit the Natural History Museum (BM) should contact the Keeper, Dep. of Botany, The Natural History Museum, Cromwell Road, London, SW7 5BD, U.K. Many visitors are expected and special arrangements have to be made to facilitate all wishes as much as possible.
Workshop
I like to draw your special attention to the workshops on large and complex families. I like to organise a Euphorbiaceae workshop on Monday or Tuesday afternoon. The workshops will take 2 hours, from 14.00--16.00 h. If possible, while further suggestions are welcome, I like to cover 5 topics during the workshop:
- Delimitation of infrafamily taxa, genera and species
- Characters
- Geographical patterns
- Local flora projects
- Computer aid
The last two topics will also be covered in special sessions or workshops, but it will be worthwhile to discuss our participation in local flora projects (Tree Flora of Sabah and Sarawak, Flora of Brunei, Flora of the Philippines, Flora of Thailand, etc.) and our use of special software (DELTA, ETI, etc.). I am planning to organise a demonstration of the ETI (Expert Center for Taxonomic Identification; see also the next newsletter) during the symposium and of our own database project LBASE, while there might be a special DELTA workshop. Especially, the DELTA and ETI software (free obtainable) will be very handy in subtracting data for local floras, if all of these floras more or less contain the same format.
The first two topics might be a bit outdated after the publication of the Missouri proceedings. However, (hopefully) many researchers will join the symposium who were absent in St. Louis and we might discuss the region specific problems, e.g. the delimitation between Cheilosa and Neoscortechinia, between Baccaurea and Aporosa, etc. Also, a discussion of the use of special character sets (e.g., stigma structures, DNA sequences) might be an important topic.
A discussion of biogeographical patterns might help to interpret variation, which will be useful especially for those not familiar with the Malesian region.
I would like to see different chairpersons for the various topics. And I like to invite you to join the symposium and especially the workshop and to volunteer yourself as a possible chairperson. Perhaps I can already stimulate potential candidates:
Delimitation: Grady Webster?
Special characters: Max van Balgooy, Kochummen, Anne Schot, Wurdack?
Geographical patterns: Max van Balgooy, Tim Whitmore, myself?
Local flora projects: Lolita Bulalaco, Kochummen, Enki Soepadmo?
Computer aid: Jeremy Bruhl, myself?
Registration
Join the symposium and register before 28th February 1995, otherwise you are charged extra. The symposium costs are £ 90.00 for full members (£ 40 for students, and £ 50 for accompanying persons), a stay in the St. Mary's University College ranges between £ 33.70 and £ 38.50 per night (including dinner and breakfast), the symposium dinner is £ 30 per person and the Wednesday excursion is £ 15 or £ 20 per person.
Changes in Malesian Euphorbiaceae Genus list
Due to Grady Webster's overview of the genera of Euphorbiaceae several changes have to be made in the list with the Malesian genera. These changes are made in the list below and typed in bold.
The genus Chorisandrachne has been added to Leptopus.
The genus Coccoceras has been added (split of from Mallotus, delimitation uncertain).
The genus Deutzianthus (including the genus Loerzingia) has been added.
Homalanthus has become Omalanthus (already indicated by Esser).
Octospermum has been added.
Plukenetia is added with the genus Pterococcus in its synonymy. Armbruster and Gillespie have likewise been moved as collaberators for Pterococcus.
The Malagasian genus Securinega has been excluded from the list; it has often been confused with Flueggea..
The genus Symphyllea has been included in the list, but it might be a synonym of Epiprinus..
The genus Synostemon has been removed as synonym of Sauropus.
Genera and Species to be revised
The following list shows the Malesian Euphorbiaceae genera in alphabetical order, the number of species estimated world-wide, estimated within Malesia, presence world-wide, and the contributor. This list is the updated version from the second newsletter
Genus |
World |
Malesia |
Presence |
Contributor |
Acalypha |
430 |
25 |
Pantropical |
|
Actephila |
35 |
6 |
Indomalesia to Australia |
|
Agrostistachys |
9 |
5 |
India to W Malesia |
|
Alchornea |
70 |
3 |
Tropical |
|
Aleurites |
6 |
1 |
W Pacific |
|
Alphandia |
3 |
1 |
New Guinea, W Pacific |
Leiden students |
Annesijoa |
1 |
1 |
New Guina |
Leiden students |
Antidesma |
160 |
60 |
Old world tropics and warm areas |
|
Aporosa |
75 |
60 |
Indomalesia to Solomons |
|
Ashtonia |
2 |
2 |
Malay Penin., Borneo |
|
Austrobuxus |
17 |
1 |
Malesia to Fiji |
|
Baccaurea |
80 |
50 |
Indomalesia to W Pacific |
Leiden Ph.D. |
Baliospermum |
6 |
1 |
India to central Malesia |
Leiden students |
Bischofia |
1 |
1 |
Indomalesia |
Leiden students |
Blachia |
12 |
1 |
India to central Malesia |
|
Blumeodendron |
6 |
6 |
Andamans, Malesia |
|
Borneodendron |
1 |
1 |
N Borneo |
|
Botryophora |
0 |
1 |
SE Asia, W Malesia |
|
Breynia |
25 |
10 |
China to Australia and New Caledonia |
|
Bridelia |
60 |
15 |
Old world tropics |
|
Cephalomappa |
5 |
5 |
S China, Malesia |
Widuri |
Chondrostylis |
2 |
1 |
SE Asia, W Malesia |
|
Choriceras |
1 |
1 |
S New Guinea, Australia |
|
Chrozophora |
12 |
1 |
Mediterranean, trop. Africa to India |
Leiden students |
Cladogynos |
1 |
1 |
SE Asia, Malesia |
|
Claoxylon |
80 |
40 |
Old world tropics |
|
Cleidion |
25 |
3 |
Pantropical |
|
Cleistanthus |
130 |
40 |
Old world tropics |
|
Clonostylis |
0 |
1 |
Synonym of Spathiostemon |
|
Cnesmone |
10 |
1 |
Assam to W Malesia |
|
Coccoceras |
4 |
4 |
Burma to Borneo |
|
Codiaeum |
6 |
10 |
Malesia to Pacific |
Wiryani |
Croton |
750 |
35 |
Tropical and warm areas |
|
Dalechampia |
110 |
1 |
Warm areas, esp. America |
|
Deutzianthus |
2 |
1 |
Indochina and Sumatra |
|
Dicoelia |
3 |
1 |
W Malesia |
|
Dimorphocalyx |
12 |
5 |
Indomalesia to Australia |
|
Doroxylon |
1 |
1 |
Malesia |
Leiden students |
Drypetes |
200 |
50 |
E Asia, S Africa |
|
Elateriospermum |
1 |
1 |
S Thailand, Malay Pen. |
Kochummen |
Endospermum |
13 |
10 |
SE Asia to Fiji |
Kochummen |
Epiprinus |
6 |
1 |
Assam to W Malesia |
|
Erismanthus |
2 |
2 |
SE Asia to W Malesia |
|
Erythrococca |
30 |
1 |
Trop. & S Africa |
|
Euphorbia |
1600 |
30 |
Cosmopolitan |
|
Excoecaria |
40 |
5 |
Old world tropics |
|
Fahrenheitia |
4 |
1 |
India to central Malesia |
Kochummen |
Flueggea |
13 |
2 |
Tropical |
|
Fontainea |
2 |
1 |
New Guinea, Australia, New Caledonia |
|
Galearia |
6 |
5 |
SE Asia to Solomons |
Metilistina Sasinggla |
Glochidion |
300 |
150 |
Tropical |
|
Hevea |
9 |
1 |
Amazonia |
Leiden students |
Homonoia |
2 |
1 |
SE Asia, Malesia |
Leiden students |
Hura |
2 |
1 |
Trop. America |
|
Hymenocardia |
5 |
1 |
Africa, SE Asia to Sumatra |
|
Jatropha |
170 |
5 |
Tropical and warm areas, N America |
|
Kairothamnus |
1 |
1 |
New Guinea |
|
Koilodepas |
10 |
5 |
India to Malesia |
Ms. Muzayyinah |
Lasiococca |
3 |
1 |
E Himalayas to Malay Pen. |
|
Leptopus |
20 |
1 |
W Himalayas to Australia |
|
Macaranga |
240 |
125 |
Old world tropics |
|
Mallotus |
140 |
50 |
Old world tropics |
Mohamad |
Manihot |
98 |
3 |
Tropical and warm America |
Leiden students |
Margaritaria |
4 |
1 |
Tropical |
|
Megistostigma |
4 |
1 |
SE Asia, W Malesia |
|
Melanolepis |
2 |
1 |
SE Asia to Pacific |
|
Micrococca |
14 |
1 |
Old world tropics |
|
Microdesmis |
10 |
2 |
Trop. Africa, SE Asia, W Malesia |
Leiden students |
Moultonianthus |
1 |
1 |
Sumatra, Borneo |
Leiden students |
Neoroeptera |
1 |
1 |
NE Australia |
|
Neotrewia |
1 |
1 |
Malesia |
Leiden students |
Octospermum |
1 |
1 |
New Guinea |
|
Omalanthus |
35 |
15 |
Trop. Asia to Australia |
|
Omphalea |
20 |
3 |
Tropical |
|
Ostodes |
4 |
1 |
E Himalayas to Borneo |
|
Pachystylidium |
1 |
1 |
India to Central Malesia |
|
Pedilanthes |
14 |
1 |
N to tropical America |
|
Petalostigma |
7 |
1 |
New Guinea, Australia |
|
Phyllanthus |
600 |
80 |
Tropical and warm areas |
|
Pimelodendron |
8 |
4 |
Malesia |
Kochummen |
Plukenetia |
15 |
1 |
Africa to Malesia |
|
Ptychopyxis |
13 |
10 |
Thailand to W Malesia, E New Guinea |
Murdoyuwono |
Reutealis |
1 |
1 |
Philippines |
Leiden students |
Richeriella |
2 |
1 |
SE China to central Malesia |
Leiden students |
Ricinus |
1 |
1 |
Europe to Africa and Middle East |
Leiden students |
Sapium |
100 |
3 |
Tropical and warm areas to Patagonia |
Kochummen |
Sauropus |
40 |
5 |
SE Asia, Indomalesia |
|
Sebastiana |
100 |
2 |
Tropical and warm areas |
|
Spathiostemon |
3 |
1 |
Thailand, W Malesia, New Guinea |
Leiden students |
Stillingia |
30 |
1 |
Tropical and warm areas |
|
Strophioblachia |
2 |
1 |
SE Asia to central Malesia |
|
Sumbaviopsis |
1 |
1 |
Assam to W Malesia |
Leiden students |
Suregada |
40 |
3 |
Old world tropics |
|
Syndyophyllum |
1 |
1 |
Sumatra, Borneo, New Guinea |
Leiden students |
Synostemon |
0 |
0 |
Old world tropics |
Synonym of Sauropus |
Tapoides |
1 |
1 |
Borneo |
Leiden students |
Trewia |
2 |
1 |
Himalayas to Hainan |
Leiden students |
Trigonopleura |
1 |
1 |
W Malesia |
van Welzen, Bulalacao, On |
Trigonostemon |
45 |
15 |
Indomalesia |
|
Wetria |
1 |
1 |
SE Asia to W Malesia, New Guinea |
Leiden students |
Genus |
World-wide |
Malesia |
Distribution |
Publication |
Chaetocarpus |
11 |
1 |
Pantropical |
Rheedea, 1995? |
Cheilosa |
1 |
1 |
W Malesia |
Blumea 38 (1993) 161-166 |
Neoscortechinia |
6 |
6 |
Burma to Solomons |
Blumea, 1994? |
Additions and Changes in the mailing list
Central National Herbarium, Botanical Survey of India, P.O. Botanic Garden, Howrah 711 103, India. |
|
Darnaedi, Dedy |
Herbarium Bogoriense, Jalan Ir. H. Juanda 22, P.O. Box 110, Bogor 16122, Indonesia. Supervisor M.Sc. students in Bogor. |
Museum für Naturkunde der Humboldt-Universität zu Berlin, Institut für Spezielle Botanik und Botanische Sammlungen Arboretum, Späthstrasse 80/81, 12437 Berlin, Germany. He will treat Cleistanthus and Bridelia in Leiden (Sept.). |
|
Queensland Herbarium (BRI), Department of Environment & Heritage, Meiers Road, Indooroopilly, Queensland 4068, Australia. He will treat: Choriceras, Fontainea, Pedilanthus, and Petalostigma |
|
Museum für Naturkunde der Humboldt-Universität zu Berlin, Institut für Spezielle Botanik und Botanische Sammlungen Arboretum, Späthstrasse 80/81, 12437 Berlin, Germany. She will treat Actephila and Antidesma (latter in Kew, Sept.). |
|
Change of address: Center for Biodiversity, Illinois Natural History Survey, 607 East Peabody Drive, Champaign, Illinois 61820, USA |
|
Mogea, Dr. J.P. |
Herbarium Bogoriense, Jalan Ir. H. Juanda 22, P.O. Box 110, Bogor 16122, Indonesia. Supervisor M.Sc. students in Bogor. |
Department of Botany, School of Plant Sciences, University of Reading, Whiteknights PO Box 221, Reading RG6 2AS, England. He is treating Blumeodendron, Botryophora |
|
Rifai, Dr. Mien A. |
Herbarium Bogoriense, Jalan Ir. H. Juanda 22, P.O. Box 110, Bogor 16122, Indonesia. Supervisor M.Sc. students in Bogor. |
Rugayah, Mrs. |
Herbarium Bogoriense, Jalan Ir. H. Juanda 22, P.O. Box 110, Bogor 16122, Indonesia. Supervisor M.Sc. students in Bogor. |
Vajravelu, Dr. R. |
Arboretum, Department of Biology, University of Central Florida, Orlando, Florida 32816-0368, U.S.A. e-mail: fdrvajra@ucf1vm.cc.ucf.edu. Addition to address. |
Widjaja, Elisabeth |
Herbarium Bogoriense, Jalan Ir. H. Juanda 22, P.O. Box 110, Bogor 16122, Indonesia. Supervisor M.Sc. students in Bogor. |
Wiryani, Ms. Endah Woro |
Herbarium Bogoriense, Jalan Ir. H. Juanda 22, P.O. Box 110, Bogor 16122, Indonesia. She will revise the genus Codiaeum. |
Department of Biology, CB # 3280, Coker Hall, University of N Carolina at Chapel Hill, Chapel Hill, NC 27599-3280, U.S.A. He is interested in DNA analyses. |